Puzzle #291 - Runespoor (Hebi)

Hebi is a really strange puzzle genre with some neat deductions available. This puzzle was inspired by a combination of thinking about how strings of odd lengths can't normally be uniquely clued by their center alone (the direction is ambiguous) and solving another Hebi puzzle using a bunch of 4s. Thanks to Eric Fox and Jamie Hargrove for testing this one for me to reassure me that it's unique!

puzz.link interface

I'm not totally back to posting regular puzzles, but I do have a few more on the way, including my secret solver constructions, ending at puzzle 300.